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Course: ENY 3005, Spring 2011
School: University of Florida
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Orders Objectives After Insect you have completed this unit you should know for each order: its common name, order name, and meaning of the order name its development, or metamorphosis 3-4 facts regarding its life history and/or economic importance its wing type, leg types and mouth adaptations 1 Question 2 Introduction The answer to the question, "What is the difference between a bee and a...

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Orders Objectives After Insect you have completed this unit you should know for each order: its common name, order name, and meaning of the order name its development, or metamorphosis 3-4 facts regarding its life history and/or economic importance its wing type, leg types and mouth adaptations 1 Question 2 Introduction The answer to the question, "What is the difference between a bee and a beetle?" may seem easy. But when you really think about it, it is a challenge to describe the difference to another person. They both have wings, six legs, and antennae. You can't just say, "Well, that looks like a bee and that looks like a beetle." There is more to it than that. By the end of this unit you should be able to tell someone without hesitation the difference between these two insects as well as many others. How do you tell the difference between a bee and a beetle? scarab beetle As you progress through the unit, you will refer to the textbook for information to fill in your study guide table. The textbook has a appendix dedicated to describing the insect orders, but more detailed descriptions are scattered in textboxes throughout chapters 7-14. honey bees 3 Classification Review 4 Classification Review Diagram You should recall from previous units how organisms are organized. An organism is first classified into a Kingdom (such as Animalia, Plantae, Fungi, etc), then Phylum, Class, Order, Family and lastly Genus and species. You have already learned that Insects are in the kingdom Animalia and the phylum Arthropoda along with lobsters, spiders and crabs. You also know that insects are in a separate class called Insecta. The class Insecta is then broken down into 30 insect orders and these orders are then separated into families. For example, a scarab beetle is in the class Insecta, the order Coleoptera and the family Scarabaeidae. A bee is in the class Insecta, the order Hymenoptera, and the family Apidae. This lineage is diagramed on the next slide. 5 6 Review Terms Review Terms: Answers This unit will familiarize you with some of the major insect orders. Before you get started, see how many of these terms you know by writing their definitions on a piece of paper. These terms will help you throughout the unit and on your lab projects. 1. Anamorphic: 2. Ametabolous: 3. Hemimetabolous: 4. Holometabolous: 5. Apterygote: 6. Paleopterous: 7. Pterygote: 8. Neoptera: 9. Exopterygote: 10. Endopterygote: Anamorphic: Development in some parainsects where abdominal segments are added during the molting process. Ametabolous: An insect that undergoes slight or no metamorphosis. The young look just like the adult. They also keep molting even after becoming an adult, something the other kinds of insects don't do. Hemimetabolous: An insect that undergoes incomplete metamorphosis. The young look very similar to the adult. Cuckoo wasp 7 Review Terms Answers Continued Holometabolous: An insect that undergoes complete metamorphosis. This type of metamorphosis includes the larval stage, pupal stage and adult stage. Ametabolous egg juvenile adult Hemimetabolous egg juvenile adult Holometabolous egg larva pupa adult 8 Hexapoda Apterygote: An insect that doesn't have wings. Paleopterous: Insects having wings, but the wings cannot fold back over the insect's abdomen. Pterygote: This term refers to all winged insects. Neoptera: Having wings, but the wings have a flexon, or hinge, that allows the wing to be folded back over the abdomen. Exopterygote: Insects with wings that develop from structures called wing pads located on the outside of the growing insect. Endopterygote: Insects with wings that develop from internal wing pads present in the larvae. I hope you enjoy learning about the insect orders. Before we get started, I want to introduce you to several taxonomic groupings. Some scientists like to group things together and others like to split them. Over the next few slides, we will look at two classification schemes, one that lumps groups into three classes and one that splits them into four. Be sure to read chapter 7 in the text to see how these schemes are derived. 9 Read textbook pages 178-183. 10 vespid wasp Four Class Scheme Hexapoda Continued These lines indicate relatedness. It is like a family tree. All six-legged arthropods, hexapods, were once classified in the Class Insecta. After some debate, some scientists group hexapods into three classes, Parainsecta, Entognatha, and Insecta and some scientists split them into four classes, Protura, Collembola, Diplura and Insecta. Each of the classification schemes are based on certain similarities between the groups. We will take a look at two cladograms that show the differing opinions of how these organisms should be classified. Gullen and Cranston, 2005, pg 182 Note: Because the parainsects are so closely related to the insects, we still consider them insects in the non-technical sense of the term and encourage you to include them in your insect collection. Read textbook page 181. 11 In this cladogram, the proturans, collembolens, diplurans and insects are all considered to be separate classes. We will study each of these groups individually, but here, you can follow the lines to see how the classes are all related. The complete cladogram is on page 182 of your 12 text. Three Class Scheme Objectives Recap These lines indicate relatedness. It is like a family tree. With those terms and ideas firmly established in your mind you are now ready to learn about the insect orders. There are 30 insect orders in the current classification scheme. We wont be able to cover some of the lesser known orders in this lecture, but you can read about them in the text if you wish. You will read about each of the more common orders in the lecture and text and will complete the chart on your study guide. Just as a reminder, after you have completed this unit you should know for each order: Daly et. al, 1998, pg.306-307 This is another example of a cladogram showing relatedness between groups. You can see that the proturans and collembolens are together in the class Parainsecta and the diplurans have been separated into the class Entognatha. Well start by learning about these more ancient groups. We will begin with the proturans and collembolens, then move on to the diplurans and then into the wingless subclass of Insecta, the apterygotes. Note that these cladograms are proposed relationships that are based on current morphological and genetic data. As new technology becomes available relationships between the groups can change to reflect new 13 information. 1. its common name 2. its development, or metamorphosis 3. 2-3 facts regarding its life history and/or economic importance 4. any special adaptations of mouthparts, legs, or wings 5. the meaning of the order name 14 thrips first tail Class Parainsecta: Order Protura Proturan Walking Video Proturans are eyeless and lack antennae. They are very small, less than 2 mm, and they undergo anamorphic development. The nymphs begin with nine segments, but an adult has twelve. The forelegs are enlarged and serve as sensory organs (like antennae). They do not have tentoria or cerci. The legs are five segmented. proturan Read textbook page 183 and 231. Notice the meaning of the order name in the top right corner. 15 16 glue wedge Proturan Defensive Posture Video Class Parainsecta: Order Collembola This is another organism that is not a part of the class Insecta. Collembolans are tiny arthropods that live in the soil where they feed on fungi and decaying vegetation. Thus, they are important decomposers in the ecosystem. They are often called springtails" because they can tuck their tail (furcula) underneath them-- similar to cocking a gun-and can snap it back out "springing" them off of the ground. collembolan Read textbook pages 183, 230 and 231. 17 Fill out the table on your study guide under the heading "Insect Orders." 18 two tail Class Entognatha: Order Diplura Entognathans are a group that have enclosed mandibles. Diplurans are a part of this group. Diplurans are soil arthropods and have over 700 described species. They are eyeless, but do have antennae. Their name comes from the fact that they have two sclerotized forceps on the end of the abdomen. The size ranges from 0.5-50mm in length. Some diplurans feed on vegetation while others are predaceous. These guys have 10 body segments and continue to molt throughout their life. silverfish Thysanurans include silverfish and firebrats. Silverfish are a typical example of insects with ametabolous development. You have probably seen these primitive insects in your bathroom. They are wingless, have silver scales, compound eyes and three caudal filaments. These guys are soft bodied and need to be preserved in alcohol. They are occasional invaders of homes and prefer moist environments. http://www.earthlife.net/insects/diplura.html dipluran Notice that the ending -ura means tail. bristle tail Class Insecta: Subclass Apterygota: Order Thysanura (Zygentoma) Read textbook pages 183-184, 231. 19 Note: You will notice that your text includes silverfish in the order Zygentoma. Many other entomological texts have silverfish in the order Thysanura. Either term is acceptable. I will refer to them as Thysanurans as we go throughout the lecture. 20 Read textbook pages 185 and 232. Class Insecta: Subclass Pterygota: Order Ephemeroptera (Mayflies) Primitive Pterygotes Daly et. al, 1998, pg 306-307 Gullen and Cranston, 2005, pg 182 The next group we will consider are the primitive pterygotes. These insects were the first to develop wings. These wings cannot fold and must be held out away from the body. The Infraclass is Paleoptera which means ancient winged ephemera - short-lived Mayfly naiads live in water-- usually under rocks. They are herbivorous and are good indicators of water quality. The nymphs have abdominal gills and three caudal filaments. When they go through metamorphosis they will go through a subimago stage and then molt into an adult mayfly like the one pictured to the left. The triangular wings held together over the body and the three mayfly caudal filaments are two identifying We are now entering the division characteristics. Ephemeropterans like of the winged insects. Notice that their ephemeral namesake, are short the name often ends in -ptera lived. The adults emerge, mate and die which means wing. not even taking time to eat. Note: a naiad is an aquatic nymph of a hemimetabolous insect. 21 22 Read textbook pages 185 and 252. Class Insecta: Subclass Pterygota: Order Odonata (Damselflies, Dragonflies) toothed ones Odonates, like the mayflies you just read about, also have aquatic immatures. Dragonfly and damselfly naiads are easy to distinguish from one another. Damselfly nymphs have three caudal gills that are absent on dragonfly nymphs. The gills on the damselfly larvae are articulated and easily break off when the animal is disturbed. Order Odonata (Damselflies, Dragonflies) continued toothed ones compound eyes damselfly dragonfly The damselfly adult has a delicate looking body. The adult holds its wings together above the abdomen. They are considered good luck and make a heart shape when they mate. Take a look around next time you are collecting near a pond or lake. Dragonfly adults have a robust body that with the outstretched wings, resembles an airplane. They both have large compound eyes that aid them in capturing prey. These guys are predators in both the immature and adult forms. Can you tell which is which? Remember, the damselfly larva has gills. 23 Read textbook pages 186 and 253-254. Read textbook pages 186 and 253-254. 24 Infraclass Neoptera: Division Exopterygota Neopteran Phylogeny Wing pads Daly et. al, 1998, pg 306 Gullen and Cranston, 2005, pg 182 predatory hemipterans In unit two we discussed the development of neopterous wings. If you remember from that discussion, neopterous wings are foldable. The classification of neopterous insect orders, as with the insect classes, is open for some debate. Before we move on, please read about this debate in the Division Neoptera section on page 186. 25 Read textbook page 186. Order Blattodea (Cockroaches) In one classification scheme, a division of the Infraclass Neoptera is Exoterygota. Exopterygotes develop their wings on the outside of the body. These wings are seen as wing pads on the developing nymph. You can see which orders are included from the cladogram above. We will now review these orders individually. Note: These wing pads are an indication that the insect has not developed into the adult stage. This is a tip you can use while you are out collecting your insects. Mantis is the Greek word for Mantids Blatta is the Greek word for cockroach Okay, one of the first exopterygotes we are going to cover is one of my favorites cockroaches. Cockroaches are natures cleanup crew, but most people consider them to be nothing but pests. While there are a few pest species, there are many more non-pest species (about 4000 to be exact). In fact, cockroaches have contributed much to science from everything from anatomy to neurology. Cockroaches are dorso-ventrally flattened and are well adapted to their environment. The pronotum is designed to Florida woods cockroach and egg case protect its head and their cursorial legs are for running. Cockroaches have many nicknames such as palmetto, water, or croton bug. Like most insects, they come in a variety of shapes and sizes. Although they are hemimetabolous and exopterygotes, some adults lack wings. Australian cockroach Order Mantodea (Praying Mantids) Mantids are known for their legs that have the ability to strike and catch prey quickly. These raptorial legs have spines that help the predator to hold onto its prey while its chewing mouthparts dig in. Mantids are the only insects that have the ability to rotate head to look over their shoulder. This allows them to spot prey and prepare for a quick grab without moving their body. Mantids have styrofoam like egg cases and are hemimetabolous. 27 Cuban cockroach 28 equal wings Greek for Mantid and Phantom Order Isoptera (Termites) Termites are also known as white ants. They are the only hemimetabolous insect with true social behavior. These insects have a caste system that includes workers, soldiers, and reproductives. The worker is the bread winner of the family. They are nearly blind and they forage soldier termites for food by building tunnels to protect themselves. The workers are charged with protecting the colony. They have a larger head than the common workers and large strong mandibles. There is about one soldier for every 100 workers in subterranean termite colonies. Reproductives are the only termites that have wings which they lose soon after finding mate and a new place to nest. Termite colonies consist of both males and females and the king remains termite reproductive with the queen in the colony. Note: Believe it or not, termites are closely related to cockroaches. This is an order that is only found in Africa. Why then should we mention it? This order, thought to be extinct, was recently found living in South and Eastern Africa. Species in this hemimetabolous order are still being named and described. The media blitzed this discovery of such a strange critter. Read more about this find on pages 189 and 349. For more information, click on attachments above and read the Gladiators article from Scientific American. praying mantis Read textbook pages 186 and 348. Read textbook pages 188 and 236. Order Mantophasmatodea (Heel Walkers or Gladiators) 26 Gullan and Cranston 2005, pg 501 Read pages textbook 189 and 349. 29 Read textbook pages 188 and 326. 30 Order Dermaptera (Earwigs) You can tell that you have now entered the section of neopterous insects. As you can see, the wings are folded on top of each other over the abdomen and are not held out to the side or straight back. Cricket Orthopterans are an important group of plant feeding insects. The hind legs of many orthopterans are modified into saltitorial legs which are designed for jumping and many have modifications for making sound. The front wings are thickened to protect the membranous fanlike wings underneath. These guys are hemimetabolous and have chewing mouthparts. A mole cricket with fossorial front legs. A large lubber grasshopper with coloring that tells predators it's nasty to eat. skin wing straight wing Order Orthoptera (Grasshoppers, Crickets) Read textbook pages 189 and 289. Believe it or not; this earwig has wings. The wings of adult earwigs are just folded and packed away under small leathery covers. Earwigs got their name from the myth that earwigs enter the ears of sleeping humans. Although they look fierce, the abdominal pincers wont hurt. They use these modified cerci for grooming and defense. These guys live in moist mulch and may be found in rotting logs, or damp places indoors. They are generalist feeders and some species, usually wingless ones, are predatory. Earwigs are hemimetabolous and the immatures are wingless. 31 earwig From the Greek phasm or phantom Read textbook pages 190 and 290. 33 uniform or same wings (Cicadas, Leafhoppers, aphids, whiteflies, scales, etc.) Homopterans differ from hemipterans by uniformity of their wings and the origin of their piercing-sucking mouthparts. Homopteran forewings have a uniform texture unlike the halfwings of their hemipteran brothers and the winged homopterans hold their wings in a tent shape over their bodies. The proboscis is cicada opistognathous meaning that it is on the ventral posterior margin of the head capsule. The proboscis, or beak, is smaller in homopterans and they use this beak to suck out juices from spittlebug vascular plants. A common representative of this suborder is the cicada. Many of you probably collected the cicada shells as a child. The nymph lives spittlebug immature underground for as many as seventeen years feeding on roots of trees before it emerges as an adult. On the next slide, you will see some other 35 Read textbook pages 192, homopteran representatives. 292 and 259. This order is split into two subborders, the Heteroptera and the Homoptera. The Homoptera suborder used to be its own order, but now it is commonly lumped under the Order Hemiptera. Hemipterans are the true Chinch Bug bugs. They have leathery fore wings and Wheel Bug membranous hind wings. The leathery forewings are halved with the hind end being membraneous (see red outline on the right). Behind the pronotum is a triangular shield (see the yellow outline to the right). The top right side of this scutellum is where you will pin the insect for your collection. The mouths of hemipterans are adapted for the piercing and sucking of plant or animal juices A common stink bug. (Notice the red rostrum on the wheel bug). Some hemipterans have scent glands, and 34 Read textbook pages some may transmit disease. 192, 292 and 259. Suborder Homoptera Continued uniform or same wings (Cicadas, Leafhoppers, aphids, whiteflies, scales, etc.) scale insect whiteflies aphids treehopper The digestive system of most homopterans has a filtering system. This allows the insect to ingest large amounts of sap and then excrete the excess water and sugars. Ants appreciate this meal of honeydew and provide protection to the insects in exchange for this sweet meal. Many homopterans, like aphids and leaf hoppers, are important carriers of plant diseases. Others are beneficial, like the lac scale that produces shellac and the cochineal insect that produces red dye. www.craftsmidwales.co.uk/ scarf2.jpg Florida walking stick Suborder Homoptera half wings Order Hemiptera: Suborder Heteroptera (True Bugs) Order Phasmatodea (Walking sticks) Insects in this order look like twigs or branches. The males are often smaller than the females as seen in this picture of a mating pair of Florida Walking Sticks with the smaller male on the back of the female. These guys move slowly and often will sway like a branch in the wind. A few species have wings, but most are brachypterous meaning they have reduced wings. Walking sticks can have some interesting ornamentation are a favorite in insect zoos. The largest insect can reach 12 inches long. 32 Read textbook pages 190 and 235. Scarf dyed with cochineal dye Read textbook pages 2-3. 36 fringe wing Order Thysanoptera (Thrips) Some thrips are common pests of plants. They inject their mouthparts and suck the plant juices of the buds, flowers or fruits of the plant causing damage to these structures. Sometimes if you pick a plant and pull it apart you will see tiny wormylooking thrips crawling on your hand. These guys have rasping mouthparts and have been known to try rasping humans for a meal. Some species are predaceous and feed on mites. Adults can be winged or wingless. The wings are thin and are covered with a fringe of hairs. Thirps are hemimetabolous, but they may have a pseudo pupal stage with a silken cocoon. This puzzles scientists and may be an instance of convergent evolution. Lice are wingless parasites of birds and mammals. There are two suborders of lice, Anoplura the sucking lice, and Mallophaga the biting lice. Anoplurans are small and have a flat body. They have only one claw and a single tarsal segment and are mainly a pest of mammals. Mallophagans are mainly a pest of birds where they feed on feathers and skin debris. They have a larger head and two claws on each of two tarsal segments. These lice are pretty host specific and may be limited to feeding on a specific species. Lice are ectoparasites and need to remain close to the host. Therefore, they attach their eggs, or nits, to the hair or feathers of the host. thrips A chewing louse, a common pest of birds. NOTE: The term thrips is used both as the singular and plural form! Read textbook pages 191 and 291. 37 Daly et. al, 1998, pg 306-307 Gullen and Cranston, 2005, pg 182 If you remember from unit two, the fourth stage of insect evolution was neopterous with holometabolous development. Well, Endopterygota is composed of insects with that description. They have a wingless larval stage that looks different from the adult. As with most of the groups, there is some debate here too. Please read page 193 under the heading Entopterygota. Read textbook pages 193-194. stable fly If you notice the names, you will see that if the word fly is separated, then it is a Dipteran. If it is attached, it is not. Example: House fly or Butterfly head lice nits Biting lice usually dont transmit disease, but sucking lice have been proven to transmit, relapsing fever (Borellia recurrentis), epidemic typhus (Rickettsia prowazeki), and trench fever (Rickettsia quintana). Trench fever was common in WWI and was a factor in the collapse of the Russian army. 38 Read textbook pages 191 and 389 . A carrion beetle. CSI anyone? 39 two wings Order Diptera (True Flies) A sucking louse on a human's skin. sheath wing Order Coleoptera (Beetles) Infraclass Neoptera: Division Endopterygota Dipterans are the true flies. They have one set of functional forewings and the hind wings have been modified into halteres which function as a gyroscope helping to maintain balance. Flies are holometabolous and the larvae are known as maggots. Maggots are soft bodied and live in moist environments where they wont desiccate. Flies have many mouth adaptations. Most are haustellate and are liquid feeders. Some flies, like mosquitoes, have piercing-sucking mouthparts, while others, like house flies have sponging and some, like horse flies have lapping bee fly mouthparts. Flies are important in several ways, some are efficient pollinators, some damage crops and some spread disease. wingless louse Order Phthiraptera (Lice) Coleoptera is the largest order of insects (about 40% of all known animals). These guys are holometabolous and the larvae are referred to as grubs. The front wings in beetles are as hard as the rest of the exoskeleton and serve to protect the membranous hind wings that are used for flight. Beetles feed on a variety of foods including everything from plants to fungus, to rotting meat, poop, and blood. Beetles have various modifications of the pronotum, mandibles and antennae that you will read about. Wow! What a pile of dung. Can you believe the Egyptians worshiped this beetle? Read textbook pages 194-195, 295-296 and 259. tiger beetle a quick predator A scarab beetle with flared antennae to catch the scent of a mate or food. A firefly or lightning bug. Of course this is neither a fly or a bug, but a beetle. 40 membrane wing Order Hymenoptera (bees, ants, and wasps) mosquito love bugs horse fly 41 Read textbook pages 196, 257, and 391-392. Hymenopterans are social insects that are specialized for their particular niche. They are holometabolous and have mandibulate mouthparts (some, like bees have modifications where their mouth is shaped like a tongue to lap nectar from flowers). They have four wings that parasitic wasp are joined together (fore and hind) by tiny hooks called hamuli. Some wasps are herbivores and form galls, others are tiny and are parasites of other insects. Many bees and wasps are important pollinators of plants. As the bee lands on a flower to suck its sweet nectar for food, it catches pollen on its body. When it lands on another flower, this pollen will fall off thereby fertilizing the bee pollinating a flower flower. Ants are common pests around homes. Those of you in the Southeastern US, are familiar with fire ants. These were accidentally introduced into the US in the 1930s and have been quite successful. An interesting fact about hymenopterans is that females develop from fertilized eggs and males develop from unfertilized eggs. Cool huh? ant Read textbook pages 197 and 325. 42 Order Lepidoptera (Butterflies and Moths) scale wing Moth and Butterfly differences Lepidoptera is the second largest order of insects behind Coleoptera. Lepidopterans are holometabolous and the larvae are known as caterpillars. While it appears that caterpillars have many legs, there are only three true pair and 2-8 pair of fleshy prolegs. Most caterpillars are herbivorous and many can cause substantial damage to crops trees, or ornamental plants. Some caterpillars can sting because they are covered with urdicating hairs. Adult leps are well known for their scaled wings. A common site at gardens and zoos is a butterfly garden. The adult butterflies and moths feed by a siphoning tube that is coiled up for storage. Moths the antennae taper to a point they are thick bodied the wings lay against the abdomen at rest Sphinx moth Butterflies the antennae end in a club the body is more delicate the wings are held upright or out to the side at rest brushfooted butterfly UF - Butler hair wing Order Trichoptera (Caddisflies) Trichopterans are often mistaken for moths. They are related, but have some differences. They have long filiform antennae and hold their wings roof-like over their body. The wings are fringed with a row of hairs along the edge. These guys are holometabolous and the immatures are aquatic. They look like caterpillars and some make nets or cases to live in. The cases are made from pebbles in the environment, so jewelry makers have discovered they can add colored beads or pebbles to the larval habitat and have the caddisflies make them custom jewelry. 43 Caddisfly adult http://www.wildscape.com/jewelry/ Dingy purplewing with curled proboscis Read textbook pages 198-199 and 296-297. Caddisfly larval case earrings 44 Monarch caterpillar and adult wingless tube Order Siphonaptera (fleas) Fleas are laterally flattened and are secondarily wingless, meaning that they once had wings. All adults are blood feeders, hematophagous, and have piercing-sucking mouthparts. The hind legs are saltorial. Fleas are holometabolous with wormlike larvae that live off of the fecal material of the adults. The pupae reside in a cocoon made up of environmental debris. This makes home infestations difficult since the pupae may blend in with the carpet. There are over 2400 species of fleas with 3/4 of them being affiliated with rodents. Fleas are medically important not just because of their irritating bites, but because they are disease vectors. They have even been known to change history. Oriental Rat Flea: Plague vector The bubonic plague was the most common form of the Black Death. Septicemic plague is the most rare form. The mortality was close to 100% (even today there is no treatment). Symptoms are a high fever and skin turning deep shades of purple due to disseminated intravascular coagulation. Yes, all from a flea bite. flea Read textbook pages 196 and 390. 45 Read textbook pages 198 and 255-256. large wings nerve wing Order Neuroptera (Lacewings, Antlions, etc.) Neuropterans are holometabolous insects that are generally considered to be beneficial. For example, the larva and adults of lacewings feed on aphids and other small insects. You can purchase lacewing eggs or larva and release them in a garden or greenhouse to control insect pests. brown lacewing Another fun neuropteran is the antlion. Antlion larvae, also known as doodle bugs, are fascinating creatures. They dig pits in the sand and wait at the bottom for others insects (like ants) to get trapped. Neuropterans are generally weak lacewing eggs fliers. 46 Order Megaloptera (dobsonflies) Another order closely related to the neuropterans is the dobsonfly. The aquatic larva, often called hellgrammites, are often used as fish bait. The males have very large mandibles. green lacewing dobsonfly male dobsonfly Order Plecoptera: (stoneflies) antlion adult Note: The text lumps all Neuropterida together. This group includes the orders, Megaloptera, Raphidioptera, and Neuroptera. Read textbook pages 195 and For a bit of neuropteran fun, visit 47 http://www.antlionfarms.com/videos___pics. 350-351. Read textbook pages 195 and 350-351. folded wing Stoneflies are another insect with aquatic nymphs. The adults emerge at the same time and live for only a few weeks. Read textbook pages 187 and 255. 48 Review Conclusion Your mind is probably spinning with all the terms and the characteristics of each order. Do you know the difference between a bee and a beetle yet? How many wings does a fly have? During an upcoming lab or in your own collection, you should be able to look at specimens and the differences will become clearer to you. Now that you know the order characteristics, you will be an expert in no time. In review, the insect orders are grouped according to whether or not they are ametabolous, hemimetabolous or holometabolous. You will also see that the insect orders are grouped according to the absence or presence of wings and whether or not the wing has a flexon, i.e. apterygote, paleopterous or neopterous. The orders are also grouped according to how the wings develop. If the wings develop from structures on the outside of the body the insect is called an exopterygote. If the wings develop from internal wing pads, the insect is called an endopterygote. Be sure you know which orders go with each classification. wingless wasp cow killer ant Note: To review the order characteristics and check your chart from the study guide, you may refer to the appendix, A Reference Guide to Orders, on pages 499-505. 49 50
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Insect ParasitismObjectives1. Differentiate the three types of symbiosis2. Describe the different ways to classify parasites3. Discuss the costs and benefits of endo and ectoparasitism4. Differentiate between parasite and parasitoid5. Define hyper,
University of Florida - ENY - 3005
Insect Pest ManagementObjectives1.2.3.4.5.6.Define the term "Pest" and discuss how it is subjective.Describe two categories of pests.Differentiate between exponential and logistic growth.Differentiate between K and r strategists.Define EIL, ET
University of Florida - ENY - 3005
ObjectivesInsecticides1. Relate four major events in the history of Pest Control2. Describe the major types of insecticides and give an example of each3. Describe in detail how organophosphates and carbamates interferewith the normal functioning of t
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ObjectivesBiological Control1.2.3.4.5.Define biological controlDescribe the three major types of biological controlDescribe six typical augmentation productsName and describe, using a specific example, three types ofcultural control methodsExp
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Medical EntomologyObjectives1. Describe the different types of medically related effects caused by arthropods(direct and indirect)2. Define the terms associated with disease transmission3. Describe the general characteristics of the 7 diseases covere
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Unit One - Arthropod ClassificationStudy GuideLesson ObjectivesBy the end of the unit you should be able to:1) Define classification.2) Describe the hierarchy in classifiying any organism.3) Describe what Linnaeus did for classification.4) Give an
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Unit Two - Insect EvolutionStudy GuideLesson ObjectivesBy the end of the unit you should be able to:Describe the four successive stages of insect evolutionDefine ametabolous, hemimetabolous and holometabolous, and the similar termsregarding wing dev
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Unit 3 - Insect OrdersStudy GuideUnit objectives:After you have completed this unit you should know for each order:1. its common name2. its development, or metamorphosis3. 3-4 facts regarding its life history and/or economic importance4. its wing t
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Unit 4 - Integument, Development & ReproductionStudy GuideUnit objectives1. Describe the three layers of an insect's integument.2. Describe the advantages and disadvantages of an exoskeleton.3. Using the proper terms for the structures involved, expl
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Unit 5 - Maintenance and MovementStudy GuideUnit objectives:1. Describe embryonic origin of alimentary canal divisions.2. Draw and label generalized alimentary canal and describe function of eachcomponent.3. Draw and label generalized insect circula
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Unit 6 - Reception & Integration: The Nervous SystemStudy GuideUnit objectives:1. Describe the origin of the insect nervous system.2. Identify the major structures of the insect nervous system and describe their function.3. Compare and contrast the p
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Unit 7 - Getting Together & Staying AliveStudy GuideUnit objectives:1. Name and describe 5 ways insects get together to mate.2. Using an example, describe each of the different ways insects protect themselves thatwere discussed in the web lecture.3.
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Unit 8 - Insect SocialityStudy GuideUnit objectives:1. Describe the difference between subsocial, parasocial and Eusocial insect behavior.2. Describe the difference in the life histories of ants and termites.3. Define trophallaxis, pseudergate,4. Ex
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Unit 9-Adaptations to HabitatsStudy GuideUnit Objectives1.Define ecology and the related terms discussed.2.Build a simple food web using only insects for the primary, secondary and tertiary trophic levels.3.Describe the ways insects have adapted t
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Unit 10 - Insect and Plant InteractionStudy GuideUnit Objectives:1. Describe how plant and insect relationships evolved together.2. Using examples, explain how insects can protect, and help propagate plants.3. Describe the ways plants protect themsel
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Unit 11 - Insect ParasitismStudy GuideUnit objectives:1.2.3.4.5.Differentiate between the three types of symbiosis.Using examples, differentiate between parasite and parasitoid.Define gregarious, multiple and hyperparasitoidism.Using examples,
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Unit 12 Insect Pest ManagementStudy GuideUnit Objectives1.2.3.4.5.6.Define the term pest and discuss how it is subjective!Describe 2 categories of pests.Differentiate between exponential and logistic growth.Differentiate between K and r strate
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Unit 13- Biological ControlStudy GuideUnit Objectives1. Relate 4 major events in the history of pest control.2. Describe the major types of insecticides and give and example of each.3. Describe in detail how organophosphates and carbamates interfere
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Unit 14 - Biological ControlStudy GuideUnit Objectives1. Define biological control.2. Describe the three major types of biological control.3. Describe six typical augmentation products.4. Explain the different components of the mole cricket biologic
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Unit 15 - Medical EntomologyUnit Objectives1. Describe the different types of medically related problems caused by arthropods.2. Define the terms associated with disease transmission3. Describe the general characteristics of the diseases covered.-Typ
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Unit 3 of Entomology[1]Unit 3: Insect Orders.[2]Now that youve taken a look at how things are classified, and how insects evolved, were going to takea look at some of the more common insect orders. After youve completed this unit, you should know, fo
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Unit 4 in Entomology[1]Unit 4 Integument, Development and Reproduction.[2]Have you ever watched a butterfly emerge from its cocoon? How does it go from being a little worm likecaterpillar in to a beautiful butterfly with scaled wings and a different
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Unit 5 of Entomology[1]Unit 5: Insect Maintenance[2]In this unit, you'll learn how insects digest their food, breathe oxygen, and circulate their body fluid. Inunit four you were introduced to the insects reproductive system by comparing it with our
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Unit 6 in Entomology[1]Unit six. Reception and Integration: The Insect Nervous System.[2]In this unit, you'll need to describe the origin of the insect nervous system, identify the major structures ofthe insect nervous system and describe their funct
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Unit 7 in Entomology[1]For all of you fans of Survivor and Lost, this unit is for you. Unit seven, getting together and stayingalive.[2]In this unit you will need to learn to name and describe five ways insects get together to mate. Using anexample,
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Unit 8 in Entomology[1]Unit eight, Insect Sociality.[2]The objectives of this unit are to describe the characteristics of subsocial and eusocial insect behavior.We will also compare and contrast the life histories of ants and termites. We will define
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Unit 9 in Entomology[1]We've learned what insects are, how they reproduce, how they digest their food, how they move around,some of their behaviors, and now we'll learn how insects adapt to their environment. Unit nine,adaptation to habitats.[2]In t
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Unit 10 in Entomology[1]This is war. Unit 10: insect and plant interactions.[2]What does the Cold War and insect/plant interaction have in common?[3]Believe it or not, plant/insect interaction is very much like the Cold War and the accompanying arms
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Unit 11 in Entomology[1]Unit 11: Insect Parasitism[2]In this unit, well differentiate between the three types of symbiosis, describe the different ways to classifyparasites, discuss the costs and benefits of endo versus ecto parasitism, differentiate
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Unit 12 in Entomology[1]From agriculture to urban pest management, there's a whole realm of opportunities out there to controlpests. Unit 12: insect pest management.[2]The objectives of this unit are to define the term pest and discuss its subjective
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Unit 13 in Entomology[1]We briefly discussed insecticides in the unit on insect pest management. Now we'll take a closer look.Unit 13: insecticides.[2]The objectives of this unit are to relate four major events in the history of pest control, describ
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Unit 14 in Entomology[1]One principle of integrated pest management is to not only judiciously use pesticides, as we learned inthe previous unit, but also to use tactics such as biological control. Unit 14: biological control.[2]The objectives of thi
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Unit 15 in Entomology[1]Unit 15: medical entomology.[2]The objectives in this unit are to describe the different types of medically related effects caused byarthropods, both direct and indirect, to define the terms associated with disease transmissio
University of Florida - FOR - 6934
OutlineAdvanced Topics in Forest Biometrics - FOR6934Review of correlation and RegressionCorrelation Simple Linear Regression (SLR)Formulae and degrees of freedom Hypothesis tests AssumptionsMultiple Linear RegressionFormulae and df Hypothesis tests
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What is the impact of transformingvariables in regression?When units of x changedAdvanced Topics in ForestBiometrics - FOR6934Regression coefficients are applied to the transformedvalue of xNo impact on tests, fit statisticsUnits of y are changed
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Assumptions of regression, correlation,etc.Advanced Topics in ForestBiometrics - FOR6934Non-linear RegressionMeeting assumptions is anextremely importantprerequisite to data analysisW e test if the DEPENDENTdata are NORMALLYDISTRIBUTEDW e test
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OutlineAdvanced Topics in Forest Biometrics - FOR6934More Non-linear regression1. 2.3. 4.Assumptions of nonlinear regression Assessing goodness of fit in nonlinear regression Plateau models Using NLMIXED versus NLINAssumptions of nonlinear regressio
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Motivating textsAdvanced Topics in ForestBiometrics FOR6934Review of General Linear ModelsWhat is a model?A scientific model is an abstraction of realityModels can be further classified:Mathematical modelsStochastic modelsStatistical modelsSchab
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Motivating textsAdvanced Topics in ForestBiometrics FOR6934Schabenberger, O. and F.J. Pierce (2002)Contemporary Statistical Models for thePlant and Soil Sciences. CRC Press, NY,NY.Mixed Models Part ILittell, R.C., G.A. Milliken, W.W. Stroup, andR
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Motivating textsAdvanced Topics in Forest Biometrics FOR6934Mixed Models Part IISchabenberger, O. and F.J. Pierce (2002) Contemporary Statistical Models for the Plant and Soil Sciences. CRC Press, NY, NY. Littell, R.C., G.A. Milliken, W.W. Stroup, and
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What do we do next, when we find significant differences among treatments?Advanced Topics in Forest Biometrics FOR6934Mean separation proceduresBack to the plantation exampleType 3 Tests of Fixed Effects Effect DeploymentID FamilyID DeploymentI*Family
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OutlineAdvanced Topics in ForestBiometrics FOR6934Poisson RegressionGeneralized linear models (GzLMs) versusGeneral linear models (GLMs)GzLMs are statistical models that combineelements of linear and nonlinear modelsGzLMs apply if the response var
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What is Logistic Regression?Advanced Topics in Forest Biometrics FOR6934Logistic RegressionA form of regression that allows the prediction of discrete (categorical) variables using continuous and/or discrete predictors Addresses the same questions that
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What is Logistic Regression?Advanced Topics in Forest Biometrics FOR6934Logistic RegressionA form of regression that allows the prediction of discrete (categorical) variables using continuous and/or discrete predictors Addresses the same questions that
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What is Multinomial LogisticRegression?Advanced Topics in ForestBiometrics FOR6934A form of logistic regression for predicting a discrete(categorical) variable with two or more categories,using continuous and/or discrete predictorsAddresses the sam
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11/21/2007Lecture outlineAdvanced Topics in Forest Biometrics - FOR6934Introduction to statistical models for spatial data Part 1 Spatial data Stationarity Isotropy Semivariogram analysisWhy spatial statistics?A sample size of one?Spatial statisti
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12/3/2007Lecture outlineAdvanced Topics in Forest Biometrics - FOR6934Introduction to statistical models for spatial data Part 2 Classical measures Distance analysis measures Nearest neighbor methods All-distance FunctionsClassic spatial autocorre
University of Florida - MGF - 1107
Mandatory Assignment 0 Digital Note CardIm one of those instructors who does the note card thing you know, fill out information on an index card and attach apicture and hand it back to me.In this assignment you will create a digital note card (instead
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NAME: _UFID - _ _ _ _ - _ _ _ _Suggested Exercises 01 Lecture 02When turning in this assignment, this sheet must be stapled to the front with all ORGANIZED scratch work behind it.Please write your final answers on this sheet in the answer block provid
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NAME: _UFID - _ _ _ _ - _ _ _ _Suggested Exercises 02 Lecture 03When turning in this assignment, this sheet must be stapled to the front with all ORGANIZED scratch work behind it.Please write your final answers on this sheet in the answer block provid
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NAME: _UFID - _ _ _ _ - _ _ _ _Suggested Exercises 03 Lecture 04When turning in this assignment, this sheet must be stapled to the front with all ORGANIZED scratch work behind it.Please write your final answers on this sheet in the answer block provid
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NAME: _UFID - _ _ _ _ - _ _ _ _Suggested Exercises 04 Lecture 05When turning in this assignment, this sheet must be stapled to the front with all ORGANIZED scratch work behind it.Please write your final answers on this sheet in the answer block provid
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NAME: _UFID - _ _ _ _ - _ _ _ _Suggested Exercises 05 Lecture 06When turning in this assignment, this sheet must be stapled to the front with all ORGANIZED scratch work behind it.Please write your final answers on this sheet in the answer block provid
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NAME: _UFID - _ _ _ _ - _ _ _ _Suggested Exercises 06 Lecture 07When turning in this assignment, this sheet must be stapled to the front with all ORGANIZED scratch work behind it.Please write your final answers on this sheet in the answer block provid