B1510_module3_7_photosynthesis

B1510_module3_7_photosynthesis - Photosynthesis: light...

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Photosynthesis: light energy to fix CO 2 and split water; make carbohydrates and oxygen
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Photosynthesis outline Overview: role of photosynthesis in energy and biogeochemistry Light as energy and capturing light energy Light reactions and the products Without producing oxygen Splitting water to produce oxygen Carbon fixation reactions Calvin-Benson cycle Rubisco C3 vs C4
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Fig. 55.3 in Purves et al. (2004) Life: The Science of Biology Energy flow through an earth surface ecosystem
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Q: Plant a seedling in a large pot with 200 kgs of soil. Water daily for 5 years. The tree now weighs 100 kgs. The soil is carefully recovered, dried and weighed. How much will the soil weigh? A. About 200 kgs B. About 150 kgs C. About 100 kgs D. About 50 kgs E. More than 200 lbs
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Q: The biomass (dry weight) of the tree comes primarily from: a) Soil b) Water c) Air
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Photoautotrophs Light as energy source for both Chemiosmotic ATP synthesis (photophosphorylation) – Reduced electron carriers for CO 2 fixation (reduction of carbon dioxide to carbohydrate) Source of virtually all organic carbon on global surface Evolved after respiration?
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Evolutionary history of photosynthesis
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Light is energy: E = h ν Wavelength and frequency are inversely related: ν = c/ λ h is Planck's constant
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Visible light is a portion of the EM spectrum
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Photosynthesis in the 3 domains of life Archaea Bacteria Eukaryotes
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Archaeal photophosphorylation http://gcat.davidson.edu/GcatWiki/index.php/Halomicrobium_mukohataei_Genome_Fall_2009 Christian Kandt, http://moose.bio.ucalgary.ca/index.php? page=Shadows_of_the_Past
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Bacteria and chloroplasts use chlorophyll to transform light energy to chemical energy Xiong et al. 2000 Science 289:1724-1730
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Absorption spectra for chlorophylls match photosynthetic action spectrum th ed.
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Did chlorophyll in bacteria co-evolve with bacterio-rhodopsin in archaea? www.sidthomas.net/images/purple.jpg
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Q: Which of these chlorophyll molecules will have the greatest potential energy after absorption of light? a) Chlorophyll a from green plants: max absorption at 680 nm b) Bacteriochlorophyll a in purple bacteria: max absorption at 870 nm c) Bacteriochlorophyll c in green bacteria: max absorption at 740 nm d) Bacteriochlorophyll g in Heliobacteria: max absorption at 790 nm
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Chlorophylls and other pigment molecules funnel light energy to a reaction center Light-activated reaction center chlorophyll molecules are good electron donors (reducing agents)
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Major Groups of Photosynthetic Phototrophic Prokaryotes Type Bacterial Group Pigment(s) Primary Products
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B1510_module3_7_photosynthesis - Photosynthesis: light...

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