Lecture32-2009-Li

Lecture32-2009-Li - Photosynthesis Lecture 32 Chapter 19...

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Photosynthesis Lecture 32 Chapter 19 1-2 Office hours: 1:00 to 2:00 pm CMM Room 142 Huilin.Li@StonyBrook.edu
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Photosynthesis 1 Introduction of chloroplasts 2. The light reactions 3. The dark reactions
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Light is electromagnetic radiation Light exists as quanta (photons) A photon is a particle as well as a wave - simultaneously
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How much energy in one photon? E = hν = hc/λ H: Planck’s constant C: speed of light λ: wave length For a 450 nm photon E 0 = 6.626x10 -34 J s X 2.998x10 8 m/s ÷ (450X10 -9 m) 1M of photon contain energy: E = 6.022X10 23 X E 0 = 6.6X3 ÷ 4.5X10 4 J mol -1 = 4.4 X 10 4 Jmol -1 = 44 kJ mol -1 ATP = -30.5 kJ mol -1 In plants, 9 ATP are produced by 8 – 10 photons
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CO 2 + H 2 O + photons → CH 2 O + O 2 1. Light oxidizes water ( light reactions ) 1. Electrons extracted from water oxidize CO 2 ( dark reactions )
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Note similarity of chloroplasts to mitochondria Chloroplast ultrastructure: 1. outer membrane 2. intermembrane space 3. inner membrane 4. stroma (aqueous fluid) 5. thylakoid lumen (inside of thylakoid) 6. thylakoid membrane 7. granum (stack of thylakoids) 8. thylakoid (lamella) 9. starch 10. ribosome 11.plastidial DNA 12. plastoglobule (drop of lipids) Light reactions occur in thylakoid membrane of chloroplasts
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Note similarity of chlorophylls to hemes (as found in hemoglobin) But differences: Metal ions: Mg/Fe Ring #: 5/4 COOH capping: w/wo Photons are absorbed by pigment molecules Isoprenoid tails increase solubility in hydrophobic env.
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The hemes of oxidative phosphorylation in mitochondria
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Multiple pigments capture light of different wavelengths Plants are green (510 nm)
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Lecture32-2009-Li - Photosynthesis Lecture 32 Chapter 19...

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