320Fall09Lecture9 - BIO 320 Cell Biology, Fall 09 Dr....

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BIO 320 Cell Biology, Fall ‘09 Dr. Thomas Bushart Lecture 9
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Topics Eukaryotic membranes Endomembranes I Topology and equivalence I Transport into/between I Nuclear transport intro
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Eukaryotes and organelles
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Special plant organelles
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Cellular organization Specialized membranes and compartments for specific tasks I Properties of an organelle Membrane composition Internal (or maybe surface) chemistry Division of cytoplasm causes some problems I We know already how to move ions How do cells get proteins and metabolites to where they need to be?
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Internal membranes The plasma membrane is typically only a minor component of total membrane
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Cell specialization Cells tend to have the same compliments of organelles Primary functions of cells tied to organelles I Liver cells have a lot of smooth ER Detoxification purposes (chemical reactions) I Pancreatic cells have a lot of rough ER and Golgi Protein synthesis and export, respectively
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Evolution of endomembranes
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Evolution of endomembranes
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Modern prokaryotes Some prokaryotes exhibit membrane invaginations
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Endosymbiotic origins Second membrane may have derived from plasma membrane (don’t believe it?….)
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Kleptoplasty Chloroplast thievery! Several different species of sea slugs are capable of retaining chloroplasts from the algae they consume or “farming” living whole-cell colonies in their bodies Apparently multiple evolutionary instances Placida cf. dendritica From http://www.seaslugforum.net/ Elysia crispata
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Topology and equivalence Invaginations and pinching off produces equivalent spaces
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320Fall09Lecture9 - BIO 320 Cell Biology, Fall 09 Dr....

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