Lecture_13_3-4-10-PDF_40984

Lecture_13_3-4-10-PDF_40984 - Lecture 13 Photosynthesis 2 1.

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Photosynthesis 2 1. Photophosphorylation 2. Dark Reactions 3. The Calvin cycle 4. Control of the Calvin cycle 5. Photorespiration Lecture 13
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Light Light To Calvin Cycle Stroma Thylakoid membrane Electron transport chain ATP synthase Inside thylakoid 1 / 2 NADP ADP + P The light reactions in the thylakoid membrane Photophosphorylation The production of ATP using the energy of sunlight
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Comparison of the flow of H + and the orientation of the ATP synthesis in mitochondria and chloroplasts
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Photophosphorylation Process of synthesis using a proton gradient created by the energy gathered from
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Chloroplasts synthesize ATP after the imposition of a pH gradient A sudden increase on the out side to pH 8.0 generates a lot of ATP. Thylakoid membrane Chloroplasts can be incubated in pH 4.0 buffer until the inside has equilibrated. Incubation for several hours Rapid change of external pH, addition of ADP and Pi
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Mt and Ch ATP synthase (F 0 F 1 -ATPase)
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1. Both MtF 0 and CF 0 are hydrophobic transmembrane proteins that contain a proton translocating mechanism. The subunit are about the same. 2. Both MtF 1 and CF 1 are hydrophilic peripheral membrane proteins with the same subunit composition. 3. Both ATP synthases are inhibited by oligomycin. – same action mechanism Mt and Ch ATP synthase (F 0 F 1 -ATPase)
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Light reaction: ( reminder ) Energy of light is conserved as "high energy" phosphoanhydride bonds of and as reducing power of Dark reaction: Chemical reactions that convert carbon dioxide (CO 2 ) and other compounds into carbohydrates (glucose) using the and
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2-step process: “ light and dark Light reactions convert solar energy to chemical energy (light) Oxidises water Generates NADPH and ATP (and O 2 ) Calvin cycle makes sugar from CO 2 ( dark) Uses NADPH and ATP Fixes CO 2
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The dark reactions are not directly dependent on light. Reactions occur in the light. light and dark actually light-dependent (light reactions) and light-independent (dark reactions)
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Energy Transduction reactions: 1. Photoreduction ( synthesis) 2. Photophosphorylation ( synthesis) Roadmap for Photosynthesis: light reaction
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4. Produces sugar Roadmap for Photosynthesis: dark reaction
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The free energy of cleavage of phosphate bond of ATP , and reducing power of NADPH , are used to fix and reduce CO 2 to form The Calvin cycle (which is also referred to as a C3 pathway) Enzymes & intermediates of the Calvin Cycle are located in the chloroplast stroma , a compartment somewhat analogous to the mitochondrial matrix.
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The overall stoichiometry for this process is therefore: 5C 3 3C 5
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Lecture_13_3-4-10-PDF_40984 - Lecture 13 Photosynthesis 2 1.

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