are uninterpretable, because the conditions of artiFcialrearing are interfering with the normal developmentalprogram.b.If the results of the first experiments show some her-itability in the broad sense, then a further analysis ispossible. This analysis requires a genetically segregatingpopulation, made from a cross between two dialectpopulations—say, A and B. A cross between malesfrom population A and females from population B andthe reciprocal cross will give an estimate of the averagedegree of dominance of genes influencing the trait andwhether there is any sex linkage. (Remember that, inbirds, the female is the heterogametic sex.) The off-spring of this cross and all subsequent crosses mustberaised in conditions that do not confuse the learnedand the genetic components of the differences, as re-vealed in the experiments in part a. If learned effectscannot be separated out, this further genetic analysis isimpossible.
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