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Recovered%20File%201 - Lecture6 AdaptationstoShade Chocolate

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Click to edit Master subtitle style Lecture 6 Adaptations to Shade Chocolate
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Vertical structure n Canopy, Subcanopy (immature and short trees), Shrub layer, Herb layer, Surface layer
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Herb phenology n Spring ephemeral n Summer green n Evergreen n Most are perennials
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Sun leaves n Small and thick (more palisade layers) n Small surface area/weight ratio (SLA) n More chloroplasts per area n Higher light saturation point n Higher concentration of compounds  associated with PS capacity n Greater respiration costs
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Shade leaves n Broad and flat (fewer and smaller palisade  cells per unit area) n High surface/area ratio (SLA) n More chl per chloroplast; higher  chlorophyll concentration per unit fresh  weight (dark green color) and more chl b  (expands spectral region over which light  can be absorbed) n Lower concentration of compounds  associated with PS capacity = lower 
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Too much light n At light levels above the light saturation point,  plants cannot use excess photons. n Mechanisms (carotenoids) exist to dissipate the  energy created by these photons. n If these mechanisms are overwhelmed,  photosynthesis may be inhibited and tissue  damage can occur.   Photoinhibition. n Shade leaves are particularly susceptible and  require a period of time (up to an hour) before they  acclimate to higher light levels and are able 
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Sunflecks
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Sunflecks n May account for up to half of carbon gain by  understory species. n Photosynthetic induction n Changes in cell metabolism and buildup of  metabolite pools, particularly rubisco. n Increase in stomatal conductance n Slow loss of photosynthetic induction so each  sunflect allows increased efficiency in driving 
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Sun and shade species n Acclimation vs. adaptation n Sun species tend to have more sun leaves  than shade leaves n Can acclimate to moderate shade n Tend not to survive at very low light 
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Recovered%20File%201 - Lecture6 AdaptationstoShade Chocolate

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