Quiz 1, 21 October 2009
This quiz is to represent only your own work
As a UC Davis student, you are bound by an
honor code; if you can’t be trusted with that, can you be trusted in a career in, say, the health
Not only does this honor code demand personal integrity, but it also requires
you to demand integrity from your peers.
This quiz is closed book, closed note.
hand in a blue scantron with the answers to the multiple-choice section of the quiz.
Make sure that it has your name, student I.D.#, and
Test form A
” filled in.
You will also need to hand in the last two pages
of this quiz, with your name on each.
Much has been learned in the last few years about syntrophic collaborations between
microbes, and how they allow complete exploitation of energy-poor environments.
Analysis of the metabolism of these organisms has taught us interesting things about
energy generation, and analysis of the composition of these cells has taught us amazing
things about the fine structure of Bacterial and Archaeal cells.
A model system for the
study of syntrophy has been the collaboration between the fermenting Bacterium
and the methanogenic Archaeon
The genome of
has been sequenced, and is full of interesting stuff…
was first enriched by H. A. Barker in 1937 from the mud of a canal
in Delft, The Netherlands. The enrichment culture fermented ethanol to acetate, butyrate,
caproate, and methane. From the enrichment two organisms growing on ethanol were
isolated. One organism, named
, ferments ethanol and acetate to butyrate,
caproate, and H
; toward the end of growth also butanol and hexanol are formed. The
other organism, named
, fermented ethanol and CO
acetate and methane.
was later found to be a coculture of a bacterium
fermenting ethanol to acetate and H
(S-organism) and an archaeon reducing CO
to methane [
]. All three organisms came to fame,
as a model organism in the early 1950s for the study of fatty acid synthesis and fatty acid
oxidation, and the S-organism and
for the development of the concept of
interspecies hydrogen transfer and of syntrophy.”
--Proceedings of the National Academy of Sciences, 12 February 2008 (