lecture20 - BioNB 2210 Lecture 20 Oct 12 2011 Lecture 20...

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BioNB 2210: Lecture 20 Oct. 12, 2011 Lecture 20: Kin Selection Professor H. Kern Reeve Learning Objectives 1. To understand the meaning and computation of genetic relatedness. 2. To understand Hamilton’s rule, inclusive fitness, and estimation of their parameters. 3. To understand how Hamilton’s rule is tested empirically. Goal: To explain how natural selection can favor a gene that causes an altruistic act toward relatives, to describe the concept of relatedness, and to present Hamilton's rule and how it is applied to behavioral data. A. Altruism : Unselfish regard for or devotion to the welfare of others. Specifically, an altruistic behavior is one that benefits (increases the fitness of) another individual (the recipient) at some cost (fitness loss) to the altruist (the actor). Thus, altruism is an interesting evolutionary paradox. Two possible "solutions" are Kin Selection and Reciprocity. B. Kin Selection 1. W.D. Hamilton's hypothesis ( J. Theoret. Biol. 7: 1, 1964) states that: "The social behaviour of a species evolves so that in each distinct behaviour-evoking situation the individual will seem to value his neighbour's fitness against his own according to the coefficients of relationship appropriate
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lecture20 - BioNB 2210 Lecture 20 Oct 12 2011 Lecture 20...

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