Heterotrophy_et_al_Feb2010_post

Heterotrophy_et_al_Feb2010_post - LIFE STYLE OF THE SMALL...

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LIFE STYLE OF THE SMALL AND DIVERSE - HETEROTROPHY, AUTOTROPHY, PHOTOTROPHY and ANAEROBIC RESPIRATION A. Heterotrophy 1. To be discussed in the upcoming lectures vis-à-vis biodegradation e - donor e - acceptor respiration organic C O 2 anaerobic resp. organic C NO 3 - , NO 2 - , SO 4 2 - SO 3 2 - , CO 3 2- , Fe 3- , Mn 4- fermentation organic C organic C 2. Overall: Large molecules [insol] ------> Small molecules [sol] -------> CO 2 , H 2 O ATP red. power synthesis, growth
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3. Example of fermentation: glucose to lactate Δ G°' (kJ/ rk ) 0.5C 6 H 12 O 6 -------> C 3 H 3 O 3 - + H + + H 2 -56 pyruvate C 3 H 3 O 3 - + H 2 --------> C 3 H 5 O 3 - -43 0.5C 6 H 12 O 6 -------> C 3 H 5 O 3 - + H + -99 glucose lactate glucose to ethanol Δ G°' (kJ/ rk ) 0.5C 6 H 12 O 6 -------> C 3 H 3 O 3 - + H + + H 2 -56 pyruvate C 3 H 3 O 3 - + H 2 O + H 2 --------> C 2 H 6 O+ HCO 3 - -63 0.5C 6 H 12 O 6 + H 2 O -------> C 2 H 6 O + H + + HCO 3 - -119 glucose ethanol Unlike what is usually illustrated in microbiology, physiology and biochemistry books, this tells us the stoichiometry and the thermodynamics of the reaction[s]
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4. Classic pathway description of glycolysis, citric acid cycle Fig 5.14, p 119 In pathway form, this tells us stepwise sequence of rks, rk where electrons are being transferred [oxidation and reduction], energy consuming and energy generating reaction. - The amount of energy released is consistent with the amount of energy needed to generate 2 ATPs.
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Embden ‐Meyerhof pathway ( glycolysis ) Free energy of hydrolysis 1 ATP = ~ 32kJ
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6. Carbon and Energy utilization C source Energy source Heterotrophs organic C organic C -chemoorganotrophs Autotrophs inorganic CO 2
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Heterotrophy_et_al_Feb2010_post - LIFE STYLE OF THE SMALL...

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