2 Lecture 33 of them completely understood Mendel’s paper although Correns came close. It remained for W.S. Sutton to recognize, in a 1902 paper, that the association of paternal and material chromosomes in pairs and their subsequent separation during meiosis constituted the physical basis of Mendelian genetics. Sutton wrote 2 of the most important papers in cytology but never received his PhD; he left science for surgery. (Sutton was a student of E.B. Wilson whose famous work The Cell (1896) described chromosome behavior and speculated on their role in heredity). The genetic revolution had a rapid impact on plant and animal improvement. Although breeders had unconsciously been using many appropriate procedures via crossing and selection in the 19th century, the emerging science of genetics and, especially, the fusion of Mendelism and quantitative genetics, put plant and animal breeding on a f rm theoretical basis. The relation between genetics and post-Mendelian plant breeding is best exempli f ed by 2 routine breeding protocols. One is the extraction and recombination of inbreds combined with selection to produce heterozygous but homogeneous
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