Lecture17 - Vernalization is a process whereby flowering is...

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Vernalization is a process whereby flowering is promoted by a cold treatment A biennial cabbage henbane Summer annual
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Vernalization is age dependent Vernalization requires a mechanism of measuring the duration of cold exposure Vernalization can be lost as a result of exposure to devernalizing conditions Vernalization takes place at SAM SAM containing mitotically active cells containing mitotically active cells 1) local chilling. 2) grafting experiment A SAM grafted from an unvernalized plant onto one that has received cold treatment will not flower while a SAM grafted from a vernalized plant onto an unvernalized one will flower.
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Cold exposure does not necessarily cause flowering but rather renders the plant competent to do so A classical experiment done by Lang and Melchers Vernalization-induced mitotically stable acquisition of the competence to flower An EPIGENETIC switch
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FLC (FLOWERING LOCUS C) is a potent repressor of flowering As its loss-of-function mutations lead to flowering without a need of cold treatment
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There are at least 3 regulatory pathways that control the expression of the FLC gene
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Tug of War between Mutations in FRI or its two homologs lead to reduced FLC mRNA level
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Tug of War between Frigida & FCA,FVE&others
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Mutations in many components of the autonomous pathway results derepression of the FLC gene, leading to late flowering
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1. Late-flowering ecotypes:
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Lecture17 - Vernalization is a process whereby flowering is...

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