His few supporters turned out to be those he had been preparing all along: Hooker, Lyell, and, most importantly, Huxley. Huxley, who up until this point had been supportive but never yet entirely convinced, was finally firmly on Darwin's side after seeing the abundant evidence and clear argumentation. Huxley was renowned for his temper, and he soon became known as "Darwin's bulldog" for his willingness to tenaciously defend the theory of evolution by natural selection. Huxley had a chance to test his teeth in a debate that took place in 1860. Darwin's most vocal critics, aside from Sedgwick, were the primate anatomist Richard Owen and the Bishop of Oxford, Samuel Wilberforce. During the annual meeting of the British Association at Oxford, after a talk that made some reference to Darwin's ideas, Wilberforce gave a half-hour tirade against Darwinism. Huxley, who was also in the
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