The Precambrian encompassed all of early Earth history and rocks from that time preserve evidence of
the evolution of life forms on a microbial level. In biology, cell theory states that a cell is the fundamental
structural and functional unit of living matter, and that the an organism a multicellular body composed
of autonomous cells with its properties being the sum of those of its cells.
Multicellular organisms (animals and plants) do not appear in the fossil record until late in Precambrian
(Late Proterozoic) time.
The Phanerozoic Eon began after the end of the Proterozoic Eon about 540 million years ago, and marks
the change when fossil remains of multicellular organisms began to appear in great abundance in the
fossil record (discussed below).
Geologic Time
Highlights of Biological Evolution
Formation of the moon in the early Solar System
Fig. 2-15. Current thought is that the Moon formed from the debris created by the collision of a small
planet-sized object with the ancestral Earth (or Proto Earth) early in the history of the Solar System
about 4.5 billion years ago.
P
R
E
C
A
M
B
R
I
A
N
About 4.56 billion years ago
Formation of Earth and Moon within the Solar System nebula (Figure 2-
15). (This is discussed in detail in Chapter 1).
About 4 billion years ago
Evidence of earliest cell-based life of Earth (prokaryotes).

About 3 billion years ago
Evidence of photosynthesis and first eukaryotic cells capable of oxygen-
based respiration.
About 3.0 to 1.8 billion years ago
World-wide deposition of banded-iron formations fundamental
to the gradual conversion of Earth atmosphere rich in carbon dioxide (CO2) to oxygen (O2) (discussed
below). This conversion took nearly a billion years. Once there was enough free oxygen in the
atmosphere, this allowed the development of an ozone layer to protect Earth from deadly solar
ultraviolet radiation (UV). UV destroys many organic organic compounds. Without an ozone layer,
intense solar UV probably would have killed life in the shallow ocean waters.
About 1.8 billion years ago
Sexual reproduction fully established in eukaryotes. Sexual reproduction
increased the rate of mutation in species, leading to increased biodiversity.
About 1 billion years ago
Earliest evidence of multicellular organisms (metazoans). Early
multicellular organism were very primitive but diversified very quickly through geologic time.
P
H
A
N
E
R
O
Z
O
I
C
E
O
N
⬇
Cambrian Period
Beginning about 540 million years ago
The beginning of the Cambrian Period started a radiation of
species preserved in the fossil record. This is, in-part, because many organisms began to develop the first
hard skeletal material as part of defensive and functional body plans. The diversity of species preserved
in Cambrian sediments is partly because soft-bodied organisms were not preserved in Precambrian-age
sediments.

Significant changes happened in the global physical environment in Cambrian time. Formation of the
ozone layer created hospitable habitats and new space for organisms to move up and utilize shallow,
warm sea environments that followed a major transgression onto the continents. Organisms were finally


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