L567 (2010): study questions for second exam. Final set, updated 26 Nov 2010. 1. Briefly describe Stanley's hypotheses for the preponderance of cross-fertilizing plants and animals. Compare his idea with Nunney's idea. Why are these ideas considered "macr

Female choice response to artificial selection on an
exaggerated male trait in a stalk-eyed fly
GERALD S. WILKINSON
AND PAUL
R. REILLOt
Departmentof Zoology, University of Maryland, College Park, Maryland 20742, U.S.A.
SUMMARY Quantitative genetic models

Math/stats/theory background.
Name_
In the space provided, write the formulas for each of the following 1. Variance 2. Covariance 3. Regression slope 4. Heritability (narrow sense) 5. Response to selection (the breeders equation) 6. Change in the mean of

L567, Evolution, Indiana University
Prepared by C. Lively
TAYLOR AND FRANKS MODEL FOR KIN SELECTION Let y be the phenotype of our focal (or target) individual Let z be the mean phenotype in the population. Keep in mind that a change in y might also change

Lecture 22 L567
L567. The ecology of speciation Speciation. The mystery of mysteries. Questions 1. Can we explain speciation with microevolutionary forces (mutation, selection, drift), or must we appeal to "macroevolutionary forces" like species selection

Here we are interested in there trade-off between the size and number of offspring. We want to know the size of offspring that maximizes individual fitness.
DEFINING THE VARIABLES. q = fraction of resouces allocated to each offspring. qmin = the fraction

from News and Views regarding Saetre et al Nature 1998
What is the main point of the study? Is it important? Is the paper easy to read and understand? What is the point of Fig. 1? Should it be included? What is the point of Fig. 2? Should it be included?

Lecture 16 L567
L567. For next time read Zeh&Zeh on website. Sexual Selection II Darwin laid out two kinds of sexual selection (i.e., variation among individuals in mating success). 1. Intrasexual selection due to malemale combat (We won't cover this) And

L567. Lecture 13.
Last time. Evolutionary stability of sexual reproduction (i.e., when is a sexual population stable to invasion and replacement by an asexual clone?) A. the cost of producing males (the "cost of sex" Maynard Smith 1978) Aside 1: effect of

Pollinator preference and the evolution of floral traits in monkeyflowers (Mimulus)
Douglas W. Schemske* and H. D. Bradshaw, Jr.
*Department of Botany and College of Forest Resources, University of Washington, Seattle, WA 98195 Edited by Barbara Anna Scha

L567. Lecture 12.
Evolutionary stability of sex and recombination. 1. Motivation. Sex and recombination are anomalies for theory, and, from a pedagological point of view, these anomalies "capture" many of the ideas we have been studying (e.g.: mutation se

L567 Lecture 7
During the previous lecture we began thinking about Game Theory. We were thinking in terms of two strategies, A and B. One way to organize the information is to put it into a "payoff matrix "Payoff" to A B When interacting with A B E(A,A) E

L567 lecture 3
L567 lecture 3: Mutation-selection balance. The breeder's equation Population Genetics revisited. Last week we derived
( 11 + 12 )
pt+1 =
Now, we want to solve for the change in p, delta p (note: we drop the
subscripts for pt to give just

L567 Lecture 2
L567 Lecture 2. A brief history of population genetics (from Provine's book), and basic theoretical population genetics. Intuitive understanding of e = mc2, or R = h2S? Intuitive understanding of R = h2S requires unpacking h2, which require

L567 Lecture
L567 Evolutionary Ecology Lecture 1. Taken mainly from Michael Ruse (The Darwinian Revolution. Chicago Univ. Press) and Ernst Mayr (One Long Argument. Harvard Univ. Press). 1. The focus of the course is on the theoretical and conceptual under

This file shows the mean payoff as a function of the frequency of hawks. the red square shows the equilibrium freq of hawks. Note that if we start with a population of all doves (q=0), selection will drive mean fitness down V 1 Cost 1.5 qhat 0.667 p 1 0.9

POPULATION GENETICS SIMULATIONS By F. Frey and C. Lively, Indiana University How to get the program Go to http:/evolution.gs.washington.edu/popgen/popg.html (This is correct for 2008. Use a browser other than Safari). Download the appropriate version of t

q p a d
0.4 0.6 2 1
alpha(b) 1.08 alpha(B) 0.72
equal to: q*(a) +p*d mean phen equal to p*(a) + q*d mean phen weighted squared dev 0.452 0.007 1.327
To run the program, change the values for p, a, and d.
freqNum B pheno freq phen*freq BBp^2 2 2 0.360 0.72

Strategy 1 Go for the Max in good years (N=5) Strategy 2 Make fewer larger babies, to increase survival in bad years Mean number of surviving offspring Arith. Geometric mean good yr bad yr good yr bad yr mean 5 2 5 2 3.50 3.16 Strategy 1 4 3 4 3 3.50 3.46

Mean, variance, covariance and regression. The mean of a trait, z, is simply the sum of values of z in the population, divided by the number of observations, n. Let i represent anyone of the individuals in the population, so it varies from 1 to n. The mea

Local Mate Competition (LMC). Here we are interested in the question: how does the number of mates affect the proportion of resources that should be allocated by a hermaphrodite to male and females function in order to maximize individual fitness?
DEFININ

recomb sa sb z
0.25 0.99 0.99 0
<- change this value to change the recombination freq. <- change this value to change selection against the a allele <- change this value to change selection against the b allele <- change this value to 1 to get independent

recomb sa sb z
0.5 0.3 0.3 5
<- change this value to change the recombination freq. <- change this value to change selection against the a allele <- change this value to change selection against the b allele <- change this value to 1 to get independent ef